In 1540, twelve-year-old Jahan arrives in Istanbul. As an animal tamer in the sultan's menagerie, he looks after the exceptionally smart elephant Chota and befriends (and falls for) the sultan's beautiful daughter, Princess Mihrimah. A palace education leads Jahan to Mimar Sinan, the empire's chief architect, who takes Jahan under his wing as they construct (with Chota's help) some of the most magnificent buildings in history. Yet even as they build Sinan's triumphant masterpieces -- the incredible Suleymaniye and Selimiye mosques -- dangerous undercurrents begin to emerge, with jealousy erupting among Sinan's four apprentices.